Eucharideae

Eucharideae
File:Eucharis candida (14277378282).jpg
Urceolina candida
Scientific classification Edit this classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Monocots
Order:	Asparagales
Family:	Amaryllidaceae
Subfamily:	Amaryllidoideae
Tribe:	Eucharideae Hutch.
Type genus
Eucharis (now a synonym of Urceolina) Planch. & Linden
Genera
See text
Synonyms
Euchareae Traub

Eucharideae is a tribe of plants within the family Amaryllidaceae. It was augmented in 2000 by Meerow et al. following a molecular phylogenetic study that revealed that many elements of the tribe Stenomesseae segregated with it, rather than separately, and were subsequently submerged in it (although there was an initial proposal to rename this clade Stenomesseae). Further revisions were made in 2020, when three genera were merged. It forms one of the tribes of the Andean subclade of the American clade of the subfamily.

Taxonomy

In Traub's 1963 monograph on the Amaryllidaceae, he conceived of a tribe Euchareae based on the type genus, Eucharis and constituted from six genera in total. In 1996, the Müller-Doblies deemed this to be polyphyletic, and redistributed the genera over three separate tribes, retaining Eucharideae as one of those tribes, but divided into two subtribes, Eucharidinae and Hymenocallidinae with a total of seven genera. In 1998, Meerow and Snijman considered these separate tribes, retaining only four genera in their Eucharideae. In 2000, the tribe was then considerably reconstituted following a deconstruction of tribe Stenomesseae based on molecular phylogenetics, resulting in seven genera (Plagiolirion was not included). However, relationships were not fully resolved.

Phylogeny

The placement of Eucharideae within subfamily Amaryllidoideae is shown in the following cladogram:

Cladogram: Tribes of subfamily Amaryllidoideae

Subfamily Amaryllidoideae Africa Tribe Amaryllideae Africa Tribe Cyrtantheae Africa Tribe Haemantheae Australasia Tribe Calostemmateae Eurasian clade Asia Tribe Lycorideae Mediterranean Tribe Galantheae Tribe Pancratieae Tribe Narcisseae American clade Hippeastroid clade Tribe Griffineae Tribe Hippeastreae Andean clade Tribe Eustephieae Tribe Eucharideae/Stenomesseae Tribe Clinantheae Tribe Hymenocallideae

A 2020 molecular phylogenetic study showed that genera accepted in 2000 required further revision. A summary cladogram from the study combined Eucharis, Caliphruria and Urceolina in a single clade, as well as showing that two species were wrongly placed.

Eucharideae	Rauhia Plagiolirion (Plagiolirion horsmannii) Phaedranassa Eucrosia Stenomesson (including Caliphruria korsakoffii) Urceolina (including Eucharis, Caliphruria, Eucrosia dodsonii)

Genera

Genera accepted in the 2020 study are:

• Eucrosia Ker Gawl.
• Phaedranassa Herb.
• Plagiolirion Baker
• Rauhia Traub
• Stenomesson Herb.
• Urceolina Rchb. (including Eucharis and Caliphruria)

Bibliography

• In Rudall et al. (1995)
• , in Kubitzki (1998). (additional excerpts)