Simolestes

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Simolestes
Temporal range: 171.6–145.5 Ma MiddleLate Jurassic[1]
File:Simolestes.jpg
Diagram of the skull of S. vorax
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Superorder: Sauropterygia
Order: Plesiosauria
Suborder: Pliosauroidea
Family: Pliosauridae
Clade: Thalassophonea
Genus: Simolestes
Andrews, 1909
Species[1]
  • S. vorax (type)
  • S. indicus (?)
Synonyms

Simolestes (meaning "snub-nosed thief") is an extinct pliosaurid genus that lived in the Middle to Late Jurassic.[2] The type specimen, NHMUK PV R 3319 is an almost complete but crushed skeleton diagnostic to Simolestes vorax, dating back to the Callovian of the Oxford Clay formation, England. The genus might also be known from the Tithonian Bhuj Formation of India (S. indicus),[3] but the referral of this species to Simolestes is dubious.[4] S. keileni from France was moved to the new genus Lorrainosaurus in 2023.[5]

Description

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File:Simolestes size chart.png
Scale diagram, presenting the largest specimens of Simolestes and Lorrainosaurus (here as a species of the former genus)

Simolestes possessed a short, high, and wide skull which was built to resist torsional forces when hunting.

The largest specimens of S. vorax reached approximately 4.6 metres (15 ft) in length, if a head to body ratio similar to Liopleurodon is applied.[4][6]

Palaeobiology

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Like most pliosaurs, Simolestes possessed salt secreting glands, which would have enabled the animal to maintain salt balance and drink seawater.[4] Recent studies on plesiosaur locomotion indicate that Simolestes, like other plesiosaurs, possessed a unique bauplan for movement, which differs from modern organisms in similar niches.[7]

File:Simolestes skull side1.jpg
Skull specimen housed at the Peterborough Museum.

Feeding habits

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Simolestes's exact feeding habits are unclear. The current consensus, however, is that the genus was primarily teuthophagous, consuming belemnites, soft teuthoids and ammonites. It is possible Simolestes was also ecologically separated from other contemporary pliosaur genera such as Liopleurodon and Pachycostasaurus by hunting in deeper waters or at night, as modern cephalopods exhibit diurnal feeding cycles, spending daylight in deeper, safer waters, and rising at night to feed.[4]

Classification

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File:Simolestes1DB.jpg
Restoration of S. vorax

The cladogram below follows a 2011 analysis by paleontologists Hilary F. Ketchum and Roger B. J. Benson, and reduced to genera only.[8]

Pliosauroidea

References

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  3. ^ R. Lydekker. 1877. Notices of new and other Vertebrata from Indian Tertiary and Secondary rocks. Records of the Geological Survey of India 10(1):30-43
  4. ^ a b c d Noè, L. F. (2001). A taxonomic and functional study of the Callovian (Middle Jurassic) Pliosauroidea (Reptilia, Sauropterygia). Chicago
  5. ^ Godefroit, P. (1994). Simolestes keileni sp. nov., un Pliosaure (Plesiosauria, Reptilia) du Bajocien supérieur de Lorraine (France). Bulletin des Académie et Société Lorraines des sciences, Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value)., 1994, tome 33, n°2, p. 77-95. 33. .
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