Coordinates: 32°15′N 4°30′W / 32.25°N 4.50°W / 32.25; -4.50

Agoudim Formation

From Wikipedia, the free encyclopedia
Jump to navigation Jump to search
Agoudim Formation
Stratigraphic range: Late ToarcianBajocian 180–168 Ma
File:Creek running by Agoudim village.jpg
Agoudim village
TypeGeological Formation
Underlies
Overlies
Thickness2,000 m (6,600 ft) (up to 5,000 m (16,000 ft) reported)
Lithology
PrimaryMarl, Marly Limestone, Bioclastic limestone
Location
LocationEr-Rich
CoordinatesLua error in Module:Coordinates at line 489: attempt to index field 'wikibase' (a nil value).
Approximate paleocoordinatesLua error: callParserFunction: function "#coordinates" was not found.
RegionHigh Atlas and Middle Atlas
CountryMorocco
Type section
Named forAgoudim
Named byM. Studer
Year defined1980
RegionEr-Rich, Midelt Province
Thickness at type section~4,500–5,000 m (14,800–16,400 ft)

The Agoudim Formation (also known as the "Marnes et Marno-Calcaires d'Agoudim") is a geological formation of late Toarcian to Bajocian (LowerMiddle Jurassic[1]) age in the central High Atlas and Middle Atlas (particularly around Rich and extending to areas like Midelt, Boulemane, and Beni Mellal) of Morocco.[2][3] The formation is also known as the "Formation Marneuse d’Amane-Ilila", "Marnes de Boulemane", "Marnes de Taffert" and "Calcaires à Cancellophycus".[2][4]

Description

[edit | edit source]
File:Location of the study area in the framework of NW Africa.png
Location of the High Atlas basin and Geological overview of the main sectors

The Agoudim Formation is a substantial geological unit primarily composed of marls and marly limestones, representing a continuation from the underlying Tagoudite or Jbel Ouchbis formations and marking a significant shift toward more clay-rich (argillaceous) sedimentation.[2][5] This shift is periodically interrupted by layers of bioclastic material (deposits rich in shell fragments and other organic debris) that increase in both frequency and thickness moving upward through the formation. It is stratigraphically divided into two main members: a lower marly member dominated by gray-blue marls with occasional nodular bioclastic limestone beds, and an upper member characterized by more calcareous and bioclastic rocks with greater lithological variability.[6] The boundary between these members is often marked by a distinctive layer of organogenic turbidites, notably the "Calcaires à Cancellophycus", which forms a prominent brown cuesta and serves as an important marker horizon for geological mapping.[5][7]

The lower member consists of monotonous, micritic marls that reflect relatively quiet hemipelagic depositional conditions, where fine clay and carbonate mud settled along with benthic material.[2] This distal sedimentation was influenced by intermittent turbiditic currents transporting detrital quartz and glauconite, indicating periodic sediment input from nearby land areas and submarine slopes. In contrast, the upper member shows increasingly energetic conditions with cycles of sedimentation starting with lumachellic limestones (shell-rich limestones with oblique lamination) and ending in thick bioclastic marls.[5][6] This member records frequent synsedimentary slumps and spectacular deformation structures like ball-and-pillow folds (phacoids), which are indicative of tectonic instability and sediment loading on unstable slopes. The evolution upward culminates with a thick limestone cap dominated by oolitic limestones and bioherms (reef-like carbonate buildups that further highlight the transition to shallower marine conditions).[6][7]

Paleoenvironment

[edit | edit source]
The Agoudim Formation records multiple Paleoenvironments in a Shallowing basin, from Cold Seep deposits to shallower Reefs

The Agoudim Formation records a progressive transition from deep to shallow marine settings under the influence of rifting, subsidence, and environmental perturbations.[8] The lower marly members were deposited in hemipelagic conditions below storm wave base, with pelagic sedimentation, fine siliciclastic influx from nearby uplifts, and reworked carbonate from Liassic platforms.[9][10] Fossils such as ammonites, radiolarians, foraminifera, small bivalves, and echinoderm debris confirm a low-energy offshore environment. Rapid subsidence and sea-level rise during the Early Bajocian led to a drowning phase of the neritic carbonate factory, favoring marl sedimentation locally enriched in organic matter.[11] Within these successions, several intervals of authigenic carbonates developed, notably yellow micritic nodules, tubular and chimney-like concretions, and conical chemoherms, interpreted as the record of hydrocarbon Cold seep. Petrographic, isotopic, and sedimentological evidence indicates that the second and third authigenic horizons formed at active seep sites, with feeder-pipes, carbonate crusts, microbialites, and bioclast-rich infills, while the yellow nodules of the lower interval likely reflect organic matter remineralization without direct seepage.[12] Strongly depleted Δ13C values (down to −19‰) point to fluids rich in dissolved inorganic carbon generated by organic matter degradation, likely through bacterial sulfate reduction and anaerobic oxidation of hydrocarbons, rather than pure methane seepage. These processes created localized hardgrounds, serpulidae and oyster colonization, and chemoherm buildups on the seafloor.[12][13] As sedimentation continued, carbonate production recovered, and three reefal complexes developed, ranging from microbial-coral patch reefs to larger coral–sponge–algal bioherms, accompanied by ooid–peloid shoals and brachiopod-rich limestones.[14][15][16] These buildups evolved from small mounds to coalescing reefs, shaped by constructive growth, storm reworking, bioerosion, and micritization.[14][15][16] The upper part of the formation is dominated by shallow platform deposits with abundant brachiopods, gastropods, corals, and reefal frameworks.[14][16][17][18] Some areas, like the Agoudim valley record excepcionally preserved platform sections, showing it´s inner ramp had high-energy ooid–peloid shoals, microbial mounds, and small patch reefs and a proximal middle ramp with mixed coral–sponge–bryozoan–brachiopod assemblages in lower energy waters.[19] The distal ramp consisted of marl–limestone alternations with storm beds and oyster–coral–sponge buildups. Local syndepositional highs and terrigenous input created semi-restricted zones with fluctuating oxygenation and nutrients.[19]

Biota

[edit | edit source]

Foranimifera

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Ammobaculites[20][21] fontinensis Talghemt Ridge II-IV Tests Lituolidae
agglutinans Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Lituolidae
Bullopora[12] sp. Jebel Bou Kendill III Tests Polymorphinidae
Citharina[20] serratocostata Talghemt Ridge IV Tests Vaginulinidae
Citharina[20] sp. Talghemt Ridge I-II Tests Vaginulinidae
Eoguttulina[20][21] bilocularis Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Polymorphinidae
sp. Talghemt Ridge IV Tests Polymorphinidae
Epistommina[21] mosquensis Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II-IV Tests Epistomininae
Flabellammina[20][21] althoffi Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge III-IV Tests Lituolidae
Garantella[21] ampasindavaensis Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Epistomininae
stellata Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Epistomininae
sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Epistomininae
Haplophragmium[20][21] subaequale Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge III-IV Tests Lituolidae
aequale Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Lituolidae
Ichthyolaria[20] carinata Talghemt Ridge I Tests Ichthyolariidae
gr. major Talghemt Ridge I Tests Ichthyolariidae
sp. Talghemt Ridge I Tests Ichthyolariidae
Lenticulina[20][21] argonauta Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge IV Tests Nodosariidae
chicheryi Talghemt Ridge I-II Tests Nodosariidae
d'orbignyi Talghemt Ridge I-II Tests Nodosariidae
münsteri Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge I-II Tests Nodosariidae
quenstedti Talghemt Ridge IV Tests Nodosariidae
subalata Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge I-II Tests Nodosariidae
toarcense Talghemt Ridge I Tests Nodosariidae
polymorpha Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Nodosariidae
Parurgonina[21] sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Parurgoninidae
Planularia[20] sp. Talghemt Ridge IV Tests Vaginulinidae
Proteonina[20] gr. difflugiformis Talghemt Ridge IV Tests Hormosinidae
Protoglobigerina?[20] spp. Talghemt Ridge I-III Tests Globigerinidae
Pseudonodosaria[20] gr. pygmea Talghemt Ridge I Tests Nodosariidae
multicostata Talghemt Ridge I Tests Nodosariidae
Reinholdella[21] dreheri Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Ceratobuliminidae
Spirillina[20] orbicula Talghemt Ridge I-II Tests Spirillinidae
Textularia[20] jurassica Talghemt Ridge IV Tests Textulariidae
Trochammina[20] globigeriniformis Talghemt Ridge IV Tests Trochamminidae
Verneuilinoides[20] cf. subvitreus Talghemt Ridge I-II Tests Verneuilinidae

Ichnotaxa

[edit | edit source]
Ichnogenus Ichnospecies Locality Member Made by Images
Chondrites[22] isp. Assameur, Jebel Taferdout III-IV
File:Chondrites trace fossil - geograph.org.uk - 6982077.jpg
Example of Chondrites trace fossil
Planolites[22] isp. Assameur, Jebel Taferdout III-IV
  • Annelids
  • Bivalves
  • Arthropods
File:HortonBluffFm 001 Planolites.jpg
Example of Planolites specimens
.
Phycodes[22] isp. Assameur, Jebel Taferdout III-IV
Thalassinoides[12][23] isp. Jebel Bou Kendill III
File:Ichnofossil Thalassinoides.jpg
Thalassinoides from France
Trypanites[22] isp. Assameur, Jebel Taferdout III-IV
Zoophycos[24] (Cancellophycus) ispp. Amellago; Talghemt Ridge; Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi I-IV
File:Zoophycos2.jpg
Zoophycos fossil from Kentucky

Brachiopoda

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Acanthothiris[25] oligocantha Rich (syncline of Jebel Bou Kendill) II Shells Athyrididae
tenuispina Rich (syncline of Jebel Bou Kendill) III Shells Athyrididae
Cymatorhynchia[26] quadriplicata Jebel Assameur; Boukendil; Rich-Gourrama area IV Shells Tetrarhynchiidae
Kallirhynchia[27] concinna Amellago, Talghemt Ridge I Shells Rhynchonellidae
Loboidothyris[25] sp. Rich (syncline of Jebel Bou Kendill) II Shells Loboidothyrididae
Lophrothyris[25] contracta Rich (syncline of Jebel Bou Kendill) II Shells Terebratulidae
Monsardithyris[16][25] cortonensis Rich (syncline of Jebel Bou Kendill) III Shells Terebratulidae
ventricosa Assameur n’Ait Fergane IV Shells Terebratulidae
sp. cf. M. loubensis Bou-Ouchène (northern platform); Rich III Shells Terebratulidae
Quadratirhynchia[27] vasconcellosi Amellago I Shells Tetrarhynchiidae
Rhactorhynchia[25] distracta Rich (syncline of Jebel Bou Kendill) III Shells Rhynchonellidae
Rhynchonelloidea[25] goyi Amellago I Shells Rhynchonellidae
Rugitela[25] cf. hughesi Rich (syncline of Jebel Bou Kendill) II–III Shells Zeilleriidae
Zeilleria[16][25] truncatella Rich (syncline of Jebel Bou Kendill) III Shells Zeilleriidae
perlata Assameur n’Ait Fergane, Boukendil IV Shells Zeilleriidae

Bryozoa

[edit | edit source]
Genus Species Locality Member Material Notes Images
Berenicea[16] sp. Boukendil, Assameur IV Colonies Bereniceidae
Collapora[16] cf. straminea Boukendil IV Colonies Atactotoechidae
Entalophora[16] cf. stokesi Boukendil IV Colonies Entalophoridae

Cnidaria

[edit | edit source]
Genus Species Locality Member Material Notes Images
Acropora[16] cf.sp. Assameur n’Ait Fergane, Boukendil IV Colonies Acroporidae
File:Cherry Blossom Acropora.jpg
Extant specimen
Aggomorphastraea[28] "sp. 1" Amellago I Colonies Latomeandridae
Allocoenia[28] "sp. 1" Amellago I Colonies Actinastreidae
"sp. 2" Amellago I Colonies Actinastreidae
Ampakabastrea[28] ampakabensis Amellago I Colonies Astrocoeniidae
Cladophyllia[16] sp. Assameur n’Ait Fergane IV Colonies Caryophylliidae
Comophyllia[16] sp. Assameur n’Ait Fergane IV Colonies Comophylliidae
File:Comophyllia Fossil.png
Comophyllia Fossil
Complexastrea[28] "sp. 1" Amellago I Colonies Thecosmiliidae
Cyathocoenia[28] faugeresi Amellago I Colonies Actinastreidae
"sp. 1" Amellago I Colonies Actinastreidae
Dendrarea[16] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Dasmosmilia[28] "sp. 1" Amellago I Colonies Dermosmiliidae
File:ZooKeys - Dasmosmilia lymani.jpeg
Dasmosmilia Fossil
Dimorpharaea[16] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Edwardoseris[16] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Enallocoenia[28] "sp. 1" Amellago I Colonies Actinastreidae
Isastrea[16][28] sp. Assameur n’Ait Fergane IV Colonies Latomeandridae/Montlivaltiidae
File:Fossil Isastrea coral from Middle Jurassic, Tabas, Iran 03.jpg
Fossil Isastrea
"sp. 1" Amellago I Colonies Latomeandridae
Kobymeandra[16] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Latiastrea[28] "sp. 1" Amellago I Colonies Latomeandridae
Microphyllia[28] irregularis Amellago I Colonies Latomeandridae
Montlivaltia[16] sp. Assameur n’Ait Fergane IV Coralla Montlivaltiidae (solitary)
Polystylidium[16] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Stylosmilia[16][28] sp. Foum Zabel (southern margin) II-III Colonies Thecosmiliidae
"sp. 1" Amellago I Colonies Thecosmiliidae
Thecocyathus[28] mactrus Amellago I Solitary corallites Caryophylliidae
Vallimeandropsis[28] lineata Amellago I Colonies Scleractinia (family uncertain)

Porifera

[edit | edit source]
Genus Species Locality Member Material Notes Images
Cheilosporites[22] tirolensis Assameur, Jebel Taferdout IV Sponge body fossils Vaceletidan
spp. Assameur, Jebel Taferdout IV Sponge body fossils Vaceletidan
Cladocoropsis[16] spp. Boukendil, Assameur IV Sponge body fossils Axinellidan
File:Axinella verrucosa - Carantec.jpg
Extant relative, Axinella
Eudea[16] cf. globata Boukendil, Assameur IV Sponge body fossils Stellispongiida

Mollusks

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Actinostreon[12][15] sp. Assameur n’Ait Fergane, Boukendil IV Shells Gryphaeidae
Alocolytoceras[1] ophioneum Gheris Valley I Shells Lytoceratidae
Ambersites[1] sp. Rich region II–III Shells Hammatoceratidae
Amphitrochus[29] abbas Rich region; Outarbat III-IV Shells Nododelphinulidae
Ancolioceras[30] opalinoides n’Zala (Jbel Aouja) I–II Shells Graphoceratidae
Asthenoceras[1] sp. Rich (Concavum–Discites zones) II–III Shells Sonniniidae
Belemnopsis[31] baculiformis Iwansitn III Rostra Belemnopseidae
pougatchevskae Iwansitn III Rostra Belemnopseidae
Bositra[1] buchi Amellago; Rich I–III Shells Posidoniidae
Botula[15] sp. Boukendil area IV Shells, borings Mytilidae
Bradfordia[1] costata Rich region III Shells Oppeliidae
sp. Rich region III Shells Oppeliidae
Calabribelus[31] combemoreli Rich (Jbel Bou Kandil), Iwansitn (Middle Atlas) II–III Rostra Holcobelidae
tetramerus Rich, Iwansitn II–III Rostra Holcobelidae
Calliphylloceras[32] C. nilssoni Amellago I Shells Phylloceratidae
Catulloceras[1] sp. n’Zala (Jbel Aouja) I Shells Graphoceratidae
Chlamys[1] sp. Rich, Amellago II–III Shells Pectinidae
Cirrus[29] sp. Aberdouz III-IV Shells Cirridae
Conobelemnopsis[31] sp. A Rich, Iwansitn III Rostra Belemnopseidae
Cotteswoldia[30] sp. n’Zala (Jbel Aouja) I Shells Graphoceratidae
Ctenostreon[1] sp. Rich, Amellago II–III Shells Limidae
Cypholioceras[30] sp. n’Zala (Jbel Aouja) II Shells Graphoceratidae
Dactylioceras[20][32] (Eodactylites) mirabile Amellago I Shells Dactylioceratidae
File:Dactylioceras NT.jpg
Dactylioceras reconstruction
(Eodactylites) pseudocommune Amellago I Shells Dactylioceratidae
(Eodactylites) simplex Talghemt Ridge I Shells Dactylioceratidae
(Orthodactylites) aff. crosbeyi Amellago I Shells Dactylioceratidae
(Orthodactylites) cf. semiannulatum Amellago I Shells Dactylioceratidae
?Dicoelites[31] sp. A Iwansitn III Rostra Dicoelitidae
Discohelix[29] subaequalis Rich region; Outarbat III-IV Shells Discohelicidae
pulchrior Rich region IV Shells Discohelicidae
sp. Rich region IV Shells Discohelicidae
Docidoceras[1] wysogorskii Rich III Shells Otoitidae
Eleganticeras[32] exaratum Amellago I Shells Dactylioceratidae
sp. Amellago I Shells Dactylioceratidae
Emileia[33] (Emileia) polymera Boukendil, Assameur IV Shells Otoitidae
Eucyclus[29] ornatus Rich region IV Shells Eucyclidae
sp. Aberdouz III Shells Eucyclidae
Eucycloidea[29] (Pseudalaria?) subangulata Rich region IV Shells Eucyclidae
Eucycloidea[29] cf. etheridgii Rich region; Outarbat III-IV Shells Eucyclidae
Eudmetoceras[1] flexuosum Rich II–III Shells Hammatoceratidae
masticonense Rich III Shells Hammatoceratidae
eudmetum Rich III Shells Hammatoceratidae
sp. Rich II–III Shells Hammatoceratidae
Euaptetoceras[1] amaltheiforme Rich III Shells Hammatoceratidae
amplectens Rich III Shells Hammatoceratidae
sp. Rich II–III Shells Hammatoceratidae
Fontannesia[1] aurita Rich II Shells Sonniniidae
montillanensis Rich II Shells Sonniniidae
sp. Rich II–III Shells Sonniniidae
Grammatodon[1] sp. Rich II–III Shells Parallelodontidae
Hammatoceras[32] H. aff. insigne n’Zala (Jbel Aouja) I Shells Hammatoceratidae
Harpoceras[32] falciferum Amellago I Shells Hildoceratidae
File:Harpoceras NT.jpg
Harpoceras reconstruction
pseudoserpentinum Amellago I Shells Hildoceratidae
serpentinum Amellago I Shells Hildoceratidae
subplanatum Amellago I Shells Hildoceratidae
Hildaites[32] forte Amellago I Shells Hildoceratidae
levisoni Amellago I Shells Hildoceratidae
cf. serpentiniformis Amellago I Shells Hildoceratidae
cf. subserpentinus Amellago I Shells Hildoceratidae
striatus Amellago I Shells Hildoceratidae
wrighti Amellago I Shells Hildoceratidae
Hildoceras[20][32] bifrons Amellago; Talghemt Ridge I Shells Hildoceratidae
File:Hildoceras sublevisoni 01.JPG
Hildoceras sublevisoni
lusitanicum Amellago; Talghemt Ridge I Shells Hildoceratidae
semipolitum Amellago; Talghemt Ridge I Shells Hildoceratidae
sublevisoni Amellago; Talghemt Ridge I Shells Hildoceratidae
bonarellii Talghemt Ridge I–II Shells Hildoceratidae
sp. Amellago I Shells Hildoceratidae
Hyperlioceras[1] walkeri Rich III Shells Graphoceratidae
incisum Rich III Shells Graphoceratidae
inclusum Rich III Shells Graphoceratidae
mundum Rich III Shells Graphoceratidae
sp. Rich III Shells Graphoceratidae
Leioceras[30] cf. comptum n’Zala (Jbel Aouja) II Shells Graphoceratidae
costusum n’Zala (Jbel Aouja) II Shells Graphoceratidae
Lithophaga[16] sp. Boukendil area IV Shells, borings Mytilidae
File:Lithophaga lithophaga 001.jpg
Lithophaga specimen
Ludwigia[30] aff. haugi n’Zala (Jbel Aouja) II Shells Graphoceratidae
Lytoceras[32][34] siemensi Amellago; n’Zala I Shells Lytoceratidae
File:Lytoceras fimbriatum.JPG
Lytoceras fimbriatum
subfimbriatum n’Zala II Shells Lytoceratidae
sp. Amellago; n’Zala I–II Shells Lytoceratidae
Maconiceras[32] soloniacense Amellago I Shells Hammatoceratidae
Malladaites[1] pertinax Rich III Shells Hammatoceratidae
Nerinea[16] sp. Assameur n’Ait Fergane IV Shells Nerineidae
Nicaniella[1] sp. Rich, Amellago II–III Shells Astartidae
Palaeonucula[1] hammeri Rich, Amellago II–III Shells Nuculidae
Palaeoneilo[1] sp. Rich III Shells Malletiidae
Phylloceras[32] sp. Amellago; n’Zala I–II Shells Phylloceratidae
File:Phylloceras isotypum life restoration.png
Phylloceras reconstruction
Pholadomya[26] (Bucardiomya) bucardium Jebel Assameur; Boukendil; Rich-Gourrama area IV Shells Pholadomyidae
Planammatoceras[30] sp. n’Zala (Jbel Aouja) II Shells Hammatoceratidae
Pleurotomaria[29] armata Rich region IV Shells Pleurotomariidae
Pleydellia[30] aalensis n’Zala (Jbel Aouja) II Shells Graphoceratidae
Polyplectus[32] pluricostatum Amellago I Shells Hammatoceratidae
sp. Amellago I Shells Hammatoceratidae
Porpoceras[32] gr. vortex-verticosum Amellago; Talghemt Ridge I Shells Dactylioceratidae
Praestrigites[1] sp. Rich III Shells Strigoceratidae
Praeoppelia[1] sp. Rich III Shells Oppeliidae
Protetragonites[34] sp. n’Zala II Shells Lytoceratidae
Protoecotraustes[1] sp. Rich III Shells Haploceratidae
Rastellum[16] sp. Assameur n’Ait Fergane, Boukendil IV Shells Ostreidae
File:Rastellum carinatum fossil oyster (Upper Cretaceous; Marovoay, Madagascar) 8 (26050424446).jpg
Rastellum fossil
Rhynchocerithium[29] sp. Rich region IV Shells Procerithiidae
Riselloidea[29] sp. Outarbat III-IV Shells Eucyclidae
Sadkia[29] richensis Rich region IV Shells Eucyclidae
Saxicava[16] sp. Boukendil area IV Shells, borings Hiatellidae
Sonninia[1] dominans Rich III Shells Sonniniidae
modestum Rich III Shells Sonniniidae
nannomorpha Rich III Shells Sonniniidae
Tmetoceras[30] T. scissum n’Zala (Jbel Aouja) I–II Shells Hammatoceratidae
Toxamblyites[33] densicostatus Boukendil, Assameur IV Shells Haploceratidae
Trigonia[1] sp. Rich III Shells Trigoniidae
Trilobiticeras[1] platygaster Rich III Shells Otoitidae
trilobitoides Rich III Shells Otoitidae
sp. Rich III Shells Otoitidae
Tylostoma[1] globosum Rich III Shells Tylostomatidae
Vacekia[30] sp. n’Zala (Jbel Aouja) I–II Shells Graphoceratidae
Zurcheria[1] sp. Rich III Shells Hammatoceratidae

Arthropods

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Bairdia[20] sp. A Talghemt Ridge I–II Valves Bairdiidae
aff. hilda Talghemt Ridge IV Valves Bairdiidae
Bairdiacypris[20][21] sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge I–II Valves Bairdiacyprididae
Cytherella[20][21] aff. concentrica Talghemt Ridge IV Valves Cytherellidae
aff. fullonica Talghemt Ridge IV Valves Cytherellidae
aff. toarcensis Talghemt Ridge I–II Valves Cytherellidae
spp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge II Valves Cytherellidae
Cytherelloidea[20] gr. catenulata Talghemt Ridge IV Valves Cytherelloididae
Favreina[22] isp. Assameur, Jebel Taferdout III-IV Coprolites Decapod Coprolites (Axiidea?)
Kinkelinella[20] K. sermoisensis Talghemt Ridge I–II Valves Kinkelinellidae
gr. sermoisensis Talghemt Ridge I–II Valves Kinkelinellidae
Micropneumatocythere[21] sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge II Valves Podocopida
Monoceratina[20] aff. ventrale Talghemt Ridge IV Valves Monoceratidae
Paracypris[20] sp. Talghemt Ridge IV Valves Cytheridae
Polycope[20] sp. Talghemt Ridge IV Valves Polycopidae
Pontocyprella[20] sp. Talghemt Ridge I–II Valves Cyprididae
Procytheridea[20][21] sp. B Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge II-IV Valves Cytherideidae
Striatojonesia[20] S. striata Talghemt Ridge IV Valves Bythocytheridae

Echinoderms

[edit | edit source]
Genus Species Locality Member Material Notes Images
Cidaris[26] sp. Jebel Assameur; Boukendil; Rich-Gourrama area IV Spines & Specimens Cidaridae
File:Cidaris cidaris MHNT.jpg
Cidaris
Diplopodia[21][26] cf. morierii Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi IV Spines & Specimens Diplopodiidae
Gymnocidaris[26] pustulosa Jebel Assameur; Boukendil; Rich-Gourrama area IV Spines & Specimens Hemicidaridae
Hemicidaris[21][26] koechlini Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi IV Spines & Specimens Hemicidaridae
File:Hemicidaris intermedia Cosmocaixa cropped.jpg
Hemicidaris
Psephechinus[21][26] globosus Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi IV Spines & Specimens Stomechinidae

Annelida

[edit | edit source]
Genus Species Locality Member Material Notes Images
Nogrobs[33] moroccensis Boukendil, Assameur IV Quadrangular tubes Serpulidae
Serpula[12][16] socialis Assameur n’Ait Fergane, Boukendil; Jebel Bou Kendill (cap bed) III–IV Tubes / colonies Serpulidae
File:Serpule (Serpula vermicularis) en rade de Brest (Ifremer 00555-66731 - 43642).jpg
Serpula vermicularis
sp. Assameur n’Ait Fergane, Boukendil; Jebel Bou Kendill (cap bed) III–IV Tubes / colonies Serpulidae
Terebella[16] lapilloides Assameur n’Ait Fergane, Boukendil; Jebel Bou Kendill (cap bed) III–IV Tubes / colonies Terebellidae
File:Terebellidae (YPM IZ 080460) 002.jpeg
Terebella

Chondrichthyes

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Strophodus[35] atlasensis Talsint (eastern High Atlas) III–IV Articulated dentition (holotype PIMUZ A/I 5181) Acrodontidae
File:Strophodus tooth.png
Strophodus atlasensis

Dinoflajellates

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Kallosphaeridium[36] sp. Amellago I-II Cysts Incertade Sedis
Luehndea[36] cirilliae Amellago I Cysts Luehndeoideae
spinosa Amellago I Cysts Luehndeoideae
Mancodinium[36] semitabulatum Amellago I-III Cysts Mancodinioideae
Mendicodinium[36] microscabratum Amellago I-II Cysts Gonyaulacales
sp. 1 Amellago I-II Cysts Gonyaulacales
Moesiodinium[36] sp. Amellago I-II Cysts Heterocapsaceae
Phallocysta[36] sp. Amellago I-II Cysts Peridiniphycideae
Wallodinium[36] cf. cylindricum Amellago I-II Cysts Incertade Sedis

Algae

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Biscutum[9][37][38] depravatum Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
grande Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
Botryococcus[36] sp. Amellago I-III Miospore Botryococcaceae
Cayeuxia[16][17] sp. Assameur n’Ait Fergane; Assoul ramp IV Filamentous thalli Cyanobacteria
Carinolithus[9][37][38] superbus Amellago; Talghemt Ridge II–III Coccoliths Carinolithaceae
Cylindroporella[16] arabica Assameur n’Ait Fergane IV Thalli Dasycladaceae
Discorhabdus[9][37][38] ignotus Amellago; Talghemt Ridge II–III Coccoliths Discorhabdaceae
striatus Amellago; Talghemt Ridge II–III Coccoliths Discorhabdaceae
Garwoodia[19] sp. Amellago canyon IV Filamentous thalli Cyanobacteria
Girvanella[16] sp. Assameur n’Ait Fergane IV Filamentous thalli Cyanobacteria
Halimeda[16] sp. (cf.) Assameur n’Ait Fergane IV Thallus fragments Halimedaceae
File:Alga (Halimeda tuna), Zanzíbar, Tanzania, 2024-05-29, DD 43.jpg
Halimeda
Lotharingius[9][37][38] hauffii Amellago; Talghemt Ridge II–III Coccoliths Lotharingiaceae
Mitrolithus[9][37][38] jansae Amellago; Talghemt Ridge II–III Coccoliths Mitrolithaceae
Permocalculus[16] sp. Assameur n’Ait Fergane, Boukendil IV Encrusting thalli Gymnocodiaceae
Rivularia[19] sp. Amellago canyon IV Filamentous thalli Rivulariaceae
Sarfatiella[16] dubari Assameur n’Ait Fergane IV Thalli Dasycladaceae
Schizosphaerella[9][37][38] sp. Amellago; Talghemt Ridge II–III Coccoliths Schizosphaerellaceae
Sellinoporella[16] sp. Assameur n’Ait Fergane IV Thalli Dasycladaceae
Similiscutum[9][37][38] finchii Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
novum Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
Solenopora[16] sp. (cf.) Assameur n’Ait Fergane IV Encrusting thalli Solenoporaceae
Tasmanites[36] sp. Amellago I-III Miospore Tasmanaceae
Triscutum[9][37][38] sullivanii Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
Watznaueria[9][37][38] colacicchii Amellago; Talghemt Ridge III Coccoliths Watznaueriaceae
fossacincta Amellago; Talghemt Ridge III Coccoliths Watznaueriaceae

Plants

[edit | edit source]
Taxa Species Locality Member Material Notes Images
Alisporites[36] spp. Amellago I Pollen Corystospermaceae
Callialasporites[36] spp. Amellago I-II Pollen Araucariaceae
File:Araucaria cunninghamii cones.jpg
Callialasporites was probably related to modern plants such as Araucaria.
Classopollis[36] sp. Amellago I Pollen Hirmeriellaceae
Cyathidites[36] spp. Amellago I Spores Cyatheaceae
File:Cyathea arborea.JPG
Cyathidites was probably related to modern plants such as Cyathea.
Deltoidospora[36] spp. Amellago I Spores Pteridophyte
Kraeuselisporites[36] reissingeri Amellago I Spores Lycopodiaceae
File:Selaginella sp at Paithalmala.jpg
Kraeuselisporites was probably related to modern plants such as Selaginella.
Ischyosporites[36] variegatus Amellago I Spores Lycopodiaceae
File:Anemia phyllitidis kz05.jpg
Ischyosporites was probably related to modern plants such as Anemia.
Neoraistrickia[36] spp. Amellago I-II Spores Selaginellaceae
Quadraeculina[36] spp. Amellago I Pollen Podocarpaceae
File:Dacrycarpus dacrydioides cones2.jpg
Quadraeculina was probably related to modern plants such as Dacrycarpus.
Zonalapollenites[36] spp. Amellago I-II Pollen Araucariaceae

See also

[edit | edit source]

References

[edit | edit source]
  1. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  2. ^ a b c d Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  3. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  4. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  5. ^ a b c Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  6. ^ a b c Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  7. ^ a b Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  8. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  9. ^ a b c d e f g h i j Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  10. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  11. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  12. ^ a b c d e f Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  13. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  14. ^ a b c Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  15. ^ a b c d Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  16. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  17. ^ a b Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  18. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  19. ^ a b c d Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  20. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  21. ^ a b c d e f g h i j k l m n o p Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  22. ^ a b c d e f Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  23. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  24. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  25. ^ a b c d e f g h Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  26. ^ a b c d e f g Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  27. ^ a b Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  28. ^ a b c d e f g h i j k l m Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  29. ^ a b c d e f g h i j Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  30. ^ a b c d e f g h i Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  31. ^ a b c d Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  32. ^ a b c d e f g h i j k l Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  33. ^ a b c Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  34. ^ a b Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  35. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  36. ^ a b c d e f g h i j k l m n o p q r s Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  37. ^ a b c d e f g h i Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  38. ^ a b c d e f g h i Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).