Thermococci

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Thermococci
File:Thermococcus gammatolerans.jpg
Thermococcus gammatolerans
Scientific classification Edit this classification
Domain: Archaea
Kingdom: Methanobacteriati
Phylum: Methanobacteriota
Class: Thermococci
Zillig and Reysenbach 2002[1]
Orders[1][2]
Synonyms
  • "Methanofastidiosia" corrig. Nobu et al. 2016
  • Protoarchaea Cavalier-Smith 2002
  • "Thermococcia" Oren, Parte & Garrity 2016
File:Schematic diagram of a Thermococcus gammatolerans cell.jpg
Schematic diagram of the Thermococcus gammatolerans cell pictured above.
File:Pyrococcus furiosus.png
Pyrococcus furiosus

Thermococci is a class of microbes within the Euryarchaeota.[2]

They live in extremely hot environments, such as hydrothermal vents,[3] and their optimal growth temperatures tend to be around 75 to 85 °C.[4] Thermococcus and Pyrococcus (literally "balls of fire") are both obligately anaerobic chemoorganotrophs.

Thermococcus prefers 70–95 °C and Pyrococcus 70-100 °C. Palaeococcus helgesonii, recently discovered in the Tyrrhenian Sea, is an aerobic chemoheterotrophic that grows at temperatures of 45–85 °C with an optimal temperature of 80 °C. Thermococcus gammatolerans sp. nov. was recently discovered in the Guaymas Basin, and it grows at temperatures from 55 to 95 °C with an optimal temperature around 88 °C with an optimal pH of 6. It has pronounced radioresistance and can survive gamma radiation at 30 kGy.[5]

Thermococcus grows on organic substrates where there is a higher capacity of elemental sulfur. This archaeon mostly grows between temperatures 60–100 degrees Celsius. The average temperature where they thrive is around 85 degrees Celsius.

The DNA structure has a circular genome with around 2,353 coding sequence, and 2,306 are identified.

Taxonomy

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This class encompasses the hyperthermophilic[6][7] members of Methanobacteriota that have a sulfur-based anaerobic respiration.[7] There is only one widely recognized order within Thermococci, Thermococcales.[8] Another proposed but not yet widely accepted order is Candidatus Methanofastidiosales.[9] Thermococcales encompasses the family Thermococcaceae, which is composed of three genus: Palaeococcus, Pyrococcus and Thermococcus.[8] Palaeococcus encompasses 3 species, while Pyrococcus and Thermococcus encompass 24 and 179 species, respectively.[10]

Philogeny

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See also

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References

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Further reading

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