Rhynchobdellida

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Jawless leeches
Branchellion parkeri
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Annelida
Clade: Pleistoannelida
Clade: Sedentaria
Class: Clitellata
Subclass: Hirudinea
Infraclass: Euhirudinea
Order: Rhynchobdellida
Blanchard, 1893
Families

Rhynchobdellida (from the Greek rhynchos, mouth, and bdellein, sucking),[1] the jawless leeches[2] or freshwater leeches,[3] are an order of aquatic leeches.[4][5] Despite the common name "freshwater leeches", species are found in both sea and fresh water. They are defined by the presence of a protrusible proboscis instead of jaws, and having colourless blood.[6][7][8] They move by "inchworming"[9] and are found worldwide. The order contains 110 species, divided into 41 genera and three families.[9] Members of the order range widely in length, usually between 7 and 40 mm.[3] They are hermaphrodite.[3] The order is not monophyletic.[10]

Appearance and eating habits

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Glossiphoniidae: Smooth Turtle Leech (Placobdella parisitica) on a snapping turtle shell. (Southern United States)

Instead of jaws and teeth, Rhynchobdellidae have protrusible proboscises, which they use to penetrate the host's skin.[3][11] Mouths of Rhynchobdellidae species are small holes from which the proboscis can be protruded.[11] The proboscis then sucks out the desired bodily fluid from the host: usually blood or coelomic fluid in the case of invertebrate victims.[10]

Rhynchobdellidae are either sanguivorous or predatory.[citation needed] Sanguivorous species usually feed on a variety of other animals, using their proboscis to host's skin. They generate anticoagulants (like mosquitoes) and natural anaesthesia to help the feeding.[3] Predatory Rhynchobdellidae have digestive enzymes that help break down their prey, which usually consists of small invertebrates such as insect larvae or amphipods.[3]

Families

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Glossiphoniidae (Freshwater jawless leeches)

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Piscicolidae: Trachelobdella lubrica is a parasite of fish.

The Glossiphoniidae, the freshwater jawless leeches,[citation needed] or leaf leeches (due to their shape)[12] are freshwater leeches, flattened, and with a poorly defined anterior sucker.[3] The family Glossiphoniidae contains one of the world's largest species of leech, the giant Amazon leech, which can grow up to 45 cm in length.[13] Many species show extended parental care, keeping eggs in nests or pouches and caring for and feeding the young.[3][14] They feed on both vertebrate and invertebrate animals.[5]

Piscicolidae (Fish leeches)

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The Piscicolidae occur in both freshwater and seawater, have cylindrical bodies, and a usually well-marked, bell-shaped, anterior sucker.[3][5] They are parasites of fish.[12][5] The family was originally divided into three subfamilies based on species' pulsatile vesicles, but the subfamilies were disbanded in 2006.[5]

Ozobranchidae: Ozobranchus jantseanus (dorsal view)

Ozobranchidae (Turtle leeches)

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Ozobranchidae are primarily parasitic on marine turtles, and are sometimes merged with the Piscicolidae.[3][5]

See also

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References

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