Coordinates: 33°1′N 4°8′W / 33.017°N 4.133°W / 33.017; -4.133

Irbzer Formation

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Irbzer Formation
Stratigraphic range: Maastrichtian-Paleocene
~70–60 Ma
TypeGeological formation
AreaFes-Boulmane
Lithology
PrimaryGray mudstone
Location
CoordinatesLua error in Module:Coordinates at line 489: attempt to index field 'wikibase' (a nil value).
Approximate paleocoordinatesLua error: callParserFunction: function "#coordinates" was not found.
RegionFes-Boulmane
CountryMorocco

The Irbzer Formation is a geological formation located in Morocco in the area of Fes-Boulmane of latest Cretaceous (Maastrichtian) to Paleocene age. The lower section of this formation consists of yellowish, cross-bedded calcareous sandstone containing shark teeth, while the layer above comprises phosphatic marls with vertebrate bone beds, including dinosaur remains.[1][2]

Description

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The Irbzer Formation is 100–440 m thick, that records the transition from marginal-marine to continental environments. It is divided into three members: Lower member (20–190 m): begins with marine-influenced phosphatic sandstones, sandy limestones and marly limestones rich in marine fossils (shark teeth, mosasaurs, bivalves), then passes upward into thick white-to-varicoloured marls, often gypsiferous, with charophytes, ostracods and dinosaur eggshell fragments.[2] Middle member (2–55 m): sharp-based terrigenous interval resting on an erosive disconformity; composed of red sandstones, channelized polymictic conglomerates and siltstones with cross-bedding, ripples and bioturbation, reflecting fluvial-alluvial deposition.[2] Finally, the Upper member (25–170 m): dominantly varicoloured or gray marls with basal gypsum in many sections, thin sandstones and occasional charophyte levels; deposited in restricted lacustrine–floodplain settings.[2]

Paleoenvironment

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File:Palaeogeographic models Phosphate basins Morocco - Herbig 1991.png
Reconstructions of the "Phosphate sea" of Morocco. The Irbzer Formation area is seen at the NE

During the Maastrichtian the inner area was situated within an anoxic marine setting, as revelated by bituminous, very oily and dolomitic black shells.[3] Palynology of the underliying and coeval El Koubbat Formation has revelated an microbiota composed of Dinoflagellates, Pterospermopsis, Scolecodonts and Tasmanaceae.[4]

The formation reflects a shallow marine to lagoonal setting, transitioning from fine sands to marls and limestones. The presence of marine organisms, including bivalves and gastropods, along with phosphatic layers containing fish remains, indicates periodic marine incursions, with intermittent lagoonal or nearshore conditions.[5] Oxidation and sedimentary discontinuities suggest episodes of exposure or subaerial erosion. The overall environment was likely a shallow, low-energy, marginal marine setting with occasional deeper marine influences.[5] No planktonic foraminifera are present, and the few benthic foraminifera offer limited ecological insights. However, ostracodes, though poorly preserved, are more abundant and provide significant information about the environment. The ostracode assemblage including indicates a shallow (0–20 m), oligohaline, moderately alkaline, and relatively warm limnic environment, likely permanent and near the shoreline, as suggested by the presence of isolated dinosaur teeth and eggshell fragments in these layers.[5][6]

Marine Amniote fauna is known from coeval layers of the El Koubbat Formation, mostly Mosasaurs (Halisaurinae, Gavialimimus, Globidens, Mosasaurus, Prognathodontini) and Elasmosauridae plesiosaurs.[7]


Biota

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Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Charophytes

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Genus Species Location Material Notes Images
Feistiella[8]
  • F. sp.
Oudiksou syncline Gyrogonites A charophyte of the family Feistiellaceae
Lagynophora[9]
  • L. liburnica
Oudiksou syncline Gyrogonites A charophyte of the family Characeae
Lamprothamnium[8]
  • L. sp.
Oudiksou syncline Gyrogonites A charophyte of the family Characeae
Porochara[9]
  • P. stacheana
Oudiksou syncline Gyrogonites A charophyte of the family Characeae
Sphaerochara[8]
  • S. cf. edda
Oudiksou syncline Gyrogonites A charophyte of the family Characeae
Stephanochara[8]
  • S. sp.
Oudiksou syncline Gyrogonites A charophyte of the family Characeae

Ostracods

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Genus Species Location Material Notes Images
Brachycythere[5]
  • B. cf. gr. sapucariensis
El Koubbat Valves A marine Ostracod of the family Brachycytheridae
Clithrocytheridea[5]
  • C. senegali
El Koubbat Valves A marine Ostracod of the family Cytherideidae
Costa[5]
  • C. sp
El Koubbat Valves A marine Ostracod of the family Trachyleberididae
Cytheretta[5]
  • C.? koubbatensis
El Koubbat Valves A marine Ostracod of the family Cytherettidae
Cytherella[5]
  • C. sp
El Koubbat Valves A marine Ostracod of the family Cytherellidae
Gomphocythere[6]
  • G. achloujensis
Achlouj Valves A freshwater Ostracod of the family Limnocytheridae
Oertliella[5]
  • O. sp.
El Koubbat Valves A marine Ostracod of the family Trachyleberididae
Rehacythereis[5]
  • R. sp
El Koubbat Valves A marine Ostracod of the family Trachyleberididae
Paracandona[6]
  • P. occitanica
Achlouj Valves A freshwater Ostracod of the family Candonidae
Paracypris[5]
  • P. cf. mdaouerensis
  • P. sp.
  • P.? sp.
El Koubbat Valves A marine Ostracod of the family Paracyprididae

Echinoderms

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Genus Species Location Material Notes Images
Cretasterias[10]
  • C. reticulatus
Bakrit Thousands of specimens A sea star of the family Asteriidae
Genus Species Location Material Notes Images
Cretalamna[1]
  • C. maroccana
Oukdiksou syncline Teeth A Shark of the family Otodontidae
File:Cretalamna reconstruction.png
Cretalmana reconstruction
Dalpiazia[1]
  • D. strollleri
Oukdiksou syncline Teeth A Chondrichthian of the family Sclerorhynchidae
Ganopristis[1]
  • G. leptodoll
Oukdiksou syncline Teeth A Chondrichthian of the family Sclerorhynchidae
Serratolamna[1]
  • S. serrata
Oukdiksou syncline Teeth A Shark of the family Serratolamnidae
Squalicorax[1]
  • S. pristodontus
  • S. bassallii
Oukdiksou syncline Teeth A Shark of the family Anacoracidae
File:Squalicorax2DB.jpg
Squalicorax reconstruction

Crocodylomorpha

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Genus Species Location Material Notes Images
Krokolithidae[1]
  • Indeterminate
Oukdiksou syncline eggshells Indeterminate Crocodilian eggshells

Sauropoda

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Genus Species Location Material Notes Images
Megaloolithus[1][11]
  • M. maghrebiensis
  • M. sp.
Achlouj 2, levels 17 to 19, eggshells Eggshells referred to Titanosauridae sauropods
File:Underside of a clutch of Megaloolithus siruguei at Pinyes locality - Tremp Formation.jpg
Example of Megaloolithus
Pseudomegaloolithus[1][11]
  • P. atlasi
Achlouj 2, level 18-19 eggshells Indeterminate Sauropod eggshells

Theropoda

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Genus Species Location Material Notes Images
Pseudogeckoolithus[1][11]
  • P. tirboulensis
Achlouj 2, levels 17 to 19 eggshells Theropod eggs referred to Maniraptora
Rodolphoolithus[1][11]
  • R. arioul
Achlouj 2, levels 18 to 19 eggshells Indeterminate eggs likely laid by non-avian or avian theropods
"Paronychodon"[1]
  • "P. sp."
Oukdiksou syncline Teeth A theropod, probably related with Troodontidae or Unenlagiinae
Tipoolithus[1][11]
  • T. achloujensis
Achlouj 2, levels 18 to 19 eggshells Theropod eggs, referred to an enantiornithine bird or small Coelurosaur

References

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