Hymenasplenium
| Hymenasplenium | |
|---|---|
| |
| Hymenasplenium cheilosorum | |
| Scientific classification Edit this classification | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Division: | Polypodiophyta |
| Class: | Polypodiopsida |
| Order: | Polypodiales |
| Suborder: | Aspleniineae |
| Family: | Aspleniaceae |
| Genus: | Hymenasplenium Hayata |
| Type species | |
| Hymenasplenium unilaterale (Lam.) Hayata. 1927
| |
| Synonyms | |
| |
Hymenasplenium is one of three genera of ferns in the Aspleniaceae (spleenwort family), in the eupolypods II clade of the order Polypodiales.[1][2] The others are Hemidictyum and Asplenium. Hymenasplenium was segregated because it is a natural grouping with differing rhizome morphology – dorsiventral v. radial for the rest of Asplenium, differing chromosome count – x=39 v. x=36 for the rest of Asplenium, and a clear monophyletic grouping based on genetic analysis. It was confirmed as a sister group to Asplenium in a 2015 molecular study of the genera.[3]
List of species
[edit | edit source]As of July 2025[update], the Checklist of Ferns and Lycophytes of the World recognized the sixty-seven species and three hybrids below.[4]
- Hymenasplenium guineense (C.Presl ex Mett.) Ranil
- (N.Murak. & Hatan.) Nakaike Li Bing Zhang, W.B.Liao & K.W.Xu
- Hymenasplenium hastifolium K.W.Xu, Li Bing Zhang & W.B.Liao
- Hymenasplenium hoffmannii Hymenasplenium basiscopicum
- (Hieron.) L.Regalado & Prada Hymenasplenium hondoense
- (N.Murak. & Hatan.) Nakaike Hymenasplenium hovenkampii
- (R.C.Moran & Sundue) L.Regalado & Prada K.W.Xu & Li Bing Zhang
- Hymenasplenium ikenoi Hymenasplenium obscurum
- (Makino) Viane Hymenasplenium obtusifolium
- Hymenasplenium inthanonense Hymenasplenium cardiophyllum
- N.Murak. & J.Yokoy. Hymenasplenium oligosorum
- Hymenasplenium kenyense Hymenasplenium szechuanense
- Li Bing Zhang, K.W.Xu & Kamau (N.Murak. & R.C.Moran) L.Regalado & Prada
- Hymenasplenium andohahelense (Hance) Nakaike
- Li Bing Zhang & K.W.Xu Hymenasplenium perriei
- Hymenasplenium laetum Hymenasplenium praestans
- (Sw.) L.Regalado & Prada Hymenasplenium speluncicola
- Hymenasplenium latedens Hymenasplenium changputungense
- (Ching) Viane & S.Y.Dong Li Bing Zhang & K.W.Xu
- Hymenasplenium laterepens Hymenasplenium pubirhizoma
- N.Murak. & X.Cheng ex Y.Fen Chang & K.Hori Hymenasplenium quangnamense
- Hymenasplenium madagascariense (Ching) Viane & S.Y.Dong
- Li Bing Zhang, W.B.Liao & K.W.Xu Hymenasplenium sinense
- Hymenasplenium murakami-hatanakae Li Bing Zhang & K.W.Xu
- Nakaike Hymenasplenium quercicola
- Hymenasplenium neocaledonicum Hymenasplenium cheilosorum
- Li Bing Zhang & K.W.Xu Hymenasplenium riparium
- Hymenasplenium ngheanense Hymenasplenium sabahense
- Li Bing Zhang, K.W.Xu & N.T.Lu Hymenasplenium amoenum
- Viane (Kunze ex Mett.) Tagawa
- Li Bing Zhang & K.W.Xu Hymenasplenium obliquissimum
- (Hayata) Sugim. Hymenasplenium chingii
- (Blume) Tagawa Hymenasplenium obtusidentatum
- Y.Fen Chang & G.Cheng Zhang K.W.Xu, Li Bing Zhang & W.B.Liao
- (L.) L.Regalado & Prada Hymenasplenium ofeliae
- (Salgado) Barcelona & Pelser Hymenasplenium delitescens
- Li Bing Zhang, K.W.Xu & Lorence Hymenasplenium ortegae
- Hymenasplenium antsirananense (Maxon) L.Regalado & Prada
- Li Bing Zhang & K.W.Xu Hymenasplenium phamhoanghoi
- Li Bing Zhang, K.W.Xu & T.T.Luong Hymenasplenium denticulatum
- (Copel.) Testo, Riibe & Sundue Hymenasplenium pseudobscurum
- Viane K.W.Xu, Li Bing Zhang & W.B.Liao
- (Ching & Z.Y.Liu) K.W.Xu & Li Bing Zhang Hymenasplenium purpurascens
- (Mett.) L.Regalado & Prada Hymenasplenium distans
- Li Bing Zhang, K.W.Xu & Liang Zhang Hymenasplenium queenslandicum
- Viane Li Bing Zhang, K.W.Xu & Liang Zhang
- (Ching) Viane & S.Y.Dong Hymenasplenium repandulum
- (Kunze) L.Regalado & Prada Hymenasplenium excisum
- (Liebm.) L.Regalado & Prada Hymenasplenium rivulare
- (Fraser-Jenk.) Viane & S.Y.Dong (C.Presl) S.Linds.
- Li Bing Zhang, K.W.Xu & C.W.Chen Hymenasplenium samoaense
- Li Bing Zhang, K.W.Xu & K.R.Wood Hymenasplenium filipes
- K.W.Xu, Li Bing Zhang & W.B.Liao Hymenasplenium solomonense
- Hymenasplenium apogamum (Copel.) Sugim.
- Li Bing Zhang, K.W.Xu & H.He Hymenasplenium subnormale
- (Copel.) Nakaike Hymenasplenium furfuraceum
- (Ching) Viane & S.Y.Dong Hymenasplenium tholiforme
- Liang Zhang, W.B.Ju & K.W.Xu (Ching) Viane & S.Y.Dong
- Hymenasplenium nigricostatum Hymenasplenium kinabaluense
- Hymenasplenium trapeziforme (Wall. ex Roxb.) comb.ined., Asplenium trapeziforme Wall. ex Roxb.
- Hymenasplenium wusugongii Hymenasplenium volubile
- Hymenasplenium triquetrum (N.Murak. & R.C.Moran) L.Regalado & Prada
- (Lam.) Hayata (N.Murak. & R.C.Moran) L.Regalado & Prada
- Hymenasplenium wangpeishanii Hymenasplenium vanuatuense
- Li Bing Zhang & K.W.Xu Li Bing Zhang, W.B.Liao & K.W.Xu
- Hymenasplenium unilaterale Li Bing Zhang & K.W.Xu
- (Bail.) D.J.Ohlsen Hymenasplenium wildii
Hybrids
[edit | edit source]- Hymenasplenium × doanyense Viane
- Viane Hymenasplenium × excurum
- Hymenasplenium × incisoserratum (Rosenst.) comb.ined., Asplenium × incisoserratum (Rosenst.) N.Murak. & R.C.Moran
Phylogeny
[edit | edit source]| Phylogeny from Fern Tree of Life[5][6] | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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References
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