Helicoidea
| Helicoidea | |
|---|---|
| File:Bradybaena similaris 60096358.jpg | |
| Bradybaena similaris | |
| Scientific classification Edit this classification | |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Gastropoda |
| Order: | Stylommatophora |
| Infraorder: | Helicoidei |
| Superfamily: | Helicoidea Rafinesque, 1815 |
| Families | |
|
See text. | |
Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the order Stylommatophora.
Taxonomy
[edit | edit source]The Bouchet et al. 2017[1] nomenclator provides an up to date system of Helicoidea. The system is in some parts preliminary, as the authors relied on unpublished (as of 2023) phylogenomic study, which did not include all New World taxa. They classified Epiphragmophoridae, Helminthoglyptidae, Humboldtianidae, Monadeniidae and Xanthonychidae (provisionally also Lysinoidae and Echinichidae) as subfamilies of the last taxon, because there are no deep splits between them in the cited unpublished study (see also Koene & Schulenburg 2005[5]).
Cepolidae, Labyrinthidae and Thysanophoridae constitute the sister group of the remaining Helicoidea.[1]
- Camaenidae Pilsbry, 1895
- Canariellidae Schileyko, 1991
- Cepolidae Ihering, 1909
- Elonidae Gittenberger, 1977
- Geomitridae C. Boettger, 1909
- Helicidae Rafinesque, 1815
- Helicodontidae Kobelt, 1904
- Hygromiidae Tryon, 1866
- Labyrinthidae Borrero, Sei, Robinson & Rosenberg, 2017
- Genus: † Laxecostula Harzhauser & Neubauer, 2020
- Pleurodontidae Ihering, 1912
- Polygyridae Pilsbry, 1895
- Sphincterochilidae Zilch, 1960
- Thysanophoridae Pilsbry, 1926
- Trissexodontidae H. Nordsieck, 1987
- Xanthonychidae Strebel & Pfeffer, 1879
- Trichodiscinidae H. Nordsieck, 1987
2015 taxonomy
[edit | edit source]Razkin et al. (2015)[6] reorganized classification into monophyletic taxa according to the molecular phylogeny. This study is focus on Western Palaearctic species. The Hygromiidae s.l. family was divided into three families, Canariellidae, Geomitridae and Hygromiidae. Moreover, the family Cochlicellidae was including within the Geomitridae family as a tribe (Cochlicellini).
The classification proposed for Western Palaearctic Helicoidea is as follows:
- Bradybaenidae (now Bradybaeninae - subfamily of Camaenidae)
- Camaenidae
- Canariellidae
- Elonidae
- Geomitridae
- Geomitrinae
- Cochlicellini
- Geomitrini
- Plentuisini
- Ponentinini
- Helicellinae
- Cernuellini
- Helicellini
- Trochoideini
- Geomitrinae
- Helicodontidae
- Helicidae
- Ariantinae
- Helicinae
- Allognathini
- Helicini
- Otalini
- Thebini
- Murellinae
- Humboldtianidae
- Hygromiidae
- Ciliellinae
- Hygromiinae
- Leptaxinae
- Sphincterochilidae
- Trissexodontidae
- Gittenbergeriinae
- Trissexodontinae
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2012 taxonomy
[edit | edit source]Thompson & Naranjo-García (2012) described a new family Echinichidae and placed it to the superfamily Xanthonychoidea.[7] Therefore, the family Xanthonychidae was moved from Helicoidea to Xanthonychoidea.
2005 taxonomy
[edit | edit source]There are 19 families within the superfamily Helicoidea according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005.[8][9]
- Helicidae
- Bradybaenidae
- Camaenidae
- Cepolidae
- Cochlicellidae
- Elonidae
- Epiphragmophoridae
- Halolimnohelicidae
- Helicodontidae
- Helminthoglyptidae
- Humboldtianidae
- Hygromiidae
- Monadeniidae
- Pleurodontidae
- Polygyridae
- Sphincterochilidae
- Thysanophoridae
- Trissexodontidae
- Xanthonychidae
According to H. Nordsieck[10] the family Xanthonychidae (sensu Hausdorf & Bouchet) is probably polyphyletic (contains several different lineages) and therefore should be divided into several families.
References
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- ^ Razkin O., Gómez-Moliner B. J., Prieto C. E., Martínez-Ortí A., Arrébola J. R., Chueca L. J. & Madeira M. J. (2015). "Molecular phylogeny of the western Palearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution 83: 99–117. Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value)..
- ^ Thompson F. G. & Naranjo-García E. (2012). "Echinichidae, a new family of dart-bearing helicoid slugs from Mexico, with the description of a new genus and three new species (Gastropoda: Pulmonata: Xanthonychoidea)". Archiv für Molluskenkunde 141(2): 197–208. Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).. preview.
- ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
- ^ Poppe G. T. & Tagaro S. P. (2006). "The new classification of Gastropods according to Bouchet & Rocroi, 2005". Visaya, février 2006: 10 pp. PDF Archived 27 September 2007 at the Wayback Machine.
- ^ Higher classification of the Helicoidea and the molecular analyses of their phylogeny (by Hartmut Nordsieck) Archived 29 November 2015 at the Wayback Machine
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