Hebius

Hebius
Japanese keelback (Hebius vibakari)
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Suborder:	Serpentes
Family:	Colubridae
Subfamily:	Natricinae
Genus:	Hebius Thompson, 1913

Hebius is a genus of snakes in the family Colubridae.

Geographic range

The genus Hebius is endemic to Asia.

Taxonomy

All of the member species of the genus Hebius were formerly placed in the genus Amphiesma, but in 2014 Guo et al. placed most species of Amphiesma in the genus Hebius. They placed other species in the genus Herpetoreas, leaving Amphiesma a monotypic taxon containing only the species Amphiesma stolatum.

Species

The following 48 species are recognized as being valid.

• Hebius andreae (Ziegler & Quyet, 2006) – Andrea’s keelback
• Hebius annamensis Bourret, 1934 – Annam keelback
• Hebius arquus (David & G. Vogel, 2010)
• Hebius atemporalis (Bourret, 1934) – Tonkin keelback
• Hebius bitaeniatus (Wall, 1925) – Kutkai keelback
• Hebius boulengeri (Gressitt, 1937) – Tai-yong keelback, Boulenger’s keelback
• Hebius celebicus (W. Peters & Doria, 1878) – Sulawesi keelback
• Hebius chapaensis Bourret, 1934 – Vietnam water snake
• Hebius clerki (Wall, 1925) – Yunnan keelback
• Hebius concelarus (Malnate, 1963)
• Hebius craspedogaster (Boulenger, 1899) – Kuatun keelback
• Hebius deschauenseei (Taylor, 1934) – northern keelback, Deschauensee’s keelback
• Hebius flavifrons (Boulenger, 1887) – Sabah keelback
• Hebius frenatus (Dunn, 1923) – bridled keelback
• Hebius groundwateri (M.A. Smith, 1922) – Groundwater's keelback
• Hebius igneus David, G. Vogel, T.Q. Nguyen, Orlov, Pauwels, Teynié & Ziegler, 2021
• Hebius inas (Laidlaw, 1901) – Malayan mountain keelback
• Hebius ishigakiensis (Malnate & Munsterman, 1960) – Yaeyama keelback
• Hebius johannis (Boulenger, 1908) – Johann's keelback
• Hebius kerinciensis (David & Das, 2003)
• Hebius khasiensis (Boulenger, 1890) – Khasi Hills keelback, Khasi keelback
• Hebius lacrima Purkayastha & David, 2019 – crying keelback
• Hebius leucomystax (David, Bain, T.Q. Nguyen, Orlov, G. Vogel, Thanh & Ziegler, 2007) – white-lipped keelback
• Hebius maximus Malnate,.1962 – western China keelback
• Hebius metusia (Inger, E. Zhao, Shaffer & G. Wu, 1990) – Wa Shan keelback
• Hebius miyajimae (Maki, 1931) – Maki's keelback
• Hebius modestus (Günther, 1875) – modest keelback
• Hebius nigriventer (Wall, 1925)
• Hebius octolineatus (Boulenger, 1904) – eight-lined keelback
• Hebius optatus (Hu & E. Zhao, 1966) – Mount Omei keelback
• Hebius parallelus (Boulenger, 1890) – Yunnan keelback
• Hebius petersii (Boulenger, 1893) – Peters's keelback
• Hebius popei (Schmidt, 1925) – Pope's keelback
• Hebius pryeri (Boulenger, 1887) – Pryer's keelback
• Hebius sanguineus (Smedley, 1931) – Cameron Highlands keelback
• Hebius sangzhiensis Zhou et al., 2019 – Sangzhi keelback
• Hebius sarasinorum (Boulenger, 1896) – Sarasin’s keelback
• Hebius sarawacensis (Günther, 1872) – Sarawak keelback
• Hebius sauteri (Boulenger, 1909) – Kosempo keelback
• Hebius septemlineatus (Schmidt, 1925) – Tengchong keelback
• Hebius taronensis (M.A. Smith, 1940) – Kachin keelback
• Hebius terrakarenorum Hauser, Smits, & David, 2022
• Hebius venningi (Wall, 1910) – Chin Hills keelback
• Hebius vibakari (H. Boie, 1826) – Japanese keelback
• Hebius viperinus (Schenkel, 1901)
• Hebius weixiensis Hou, Yuan, Wei, G. Zhao, G. Liu, Y. Wu, Shen, Chen, Guo & Che, 2021 – Weixi keelback
• Hebius yanbianensis Q. Liu, Zhong, Wang, Y. Liu & Guo, 2018) – Yanbian keelback
• Hebius youjiangensis Yang, Xu, Wu, Gong, Huang & Huang, 2023

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Hebius.

Further reading

• Guo, Peng; Zhu, Fei; Liu, Qin; Zhang, Liang; Li, Jian X.; Huang, Yu Y.; Pyron, R. Alexander (2014). "A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species". Zootaxa 3873 (4): 425–440.
• Thompson JC (1913). "Contributions to the Anatomy of the Ophidia". Proceedings of the Zoological Society of London 1913: 414–425. (Hebius, new genus, p. 424).