Hadrianus (turtle)

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Hadrianus
Temporal range: 55.4–33.9 Ma Eocene[1]
"Hadrianus majusculus"
Hadrianus majusculus
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Testudines
Suborder: Cryptodira
Clade: Pantestudinoidea
Superfamily: Testudinoidea
(unranked): Pantestudinidae
Genus: Hadrianus
Cope, 1872[2][1]
Species
  • H. corsoni
  • H. majusculus

Hadrianus is an extinct genus of tortoise belonging to the Testudinidae[1] found in the United States, the Yolomécatl Formation of Mexico, the Alai Beds of Kyrgyzstan and Spain[citation needed] and believed to be the oldest true tortoise known.[3] The genus is thought to be closely related to the genus Manouria.[3] The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe.[3] Evangelos Vlachos (2018) reassessed the North American species attributed to the genus, and determined only two as accepted namely H. corsoni & H. majusculus. The remaining species were identified as either junior synonyms, moved to other genera or considered nomen dubium due to incomplete fossils.[4]

Taxonomy

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The species 'Testudo' sharanensis is hypothesized to actually be a species of Hadrianus.[5]

  • Hadrianus corsoni (Leidy, 1871)
    • Synonyms Emys carteri Leidy, 1871; Testudo hadriana Cope, 1871; Hadrianus quadratus Cope, 1871; Hadrianus octonaria Cope, 1871; Hadrianus tumidus Hay 1908; Hadrianus robustus Gilmore, 1915; Hadrianus utahensis Gilmore, 1915; Geochelone gilmorei Auffenberg, 1974
  • Hadrianus majusculus Hay, 1904

Nomen dubium

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  • "Hadrianus allabiatus" Cope, 1871

Moved species

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References

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  1. ^ a b c Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  2. ^ E. D. Cope. 1872. Second account of new Vertebrata from the Bridger Eocene. Proceedings of the American Philosophical Society (separate) 1-3
  3. ^ a b c Ehret, Dana Joseph 2004 "Skeletochronology as a method of aging Oligocene Gopherus laticuneus and Stylemys nebrascensis, using Gopherus polyphemus as a modern analog" Thesis, University of Florida.
  4. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).
  5. ^ Lua error in Module:Citation/CS1/Configuration at line 2172: attempt to index field '?' (a nil value).

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